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Carex wahuensis subsp. wahuensis

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Poales Family: Cyperaceae Genus: Carex

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Description

Key Characters:

Growth Form: Rhizomatous perennials forming large, dense clumps.

Stems: Culms trigonous, (15–)40–100 cm tall, bearing spikes ⅓–1⁄2 from base upwards.

Roots:

Leaves: Leaves simple. 3-ranked, up to 8 per culm. Blades linear, flat 1.5–2 mm wide. Blades linear, flat > 3 mm wide. Apex gradually tapering and long-acuminate. Apex gradually tapering, long-acuminate. Upper surfaces deep green; lower surfaces scabrous on veins; blades coriaceous. Margins entire. Sheathing; sheaths laxly surrounding culm bases, reddish brown to dark brown, disintegrating into parallel fibers. Stipules absent.

Flowers: Flowers in inflorescences (2–)4–9(–14)–nodose, with internodes 2–5 cm long, the spikes solitary at nodes, nodes occasionally with 2 spikes or with 3 secondary spikes, each spike pedunculate, cylindrical, 0.8–1.5 cm long, up to 5 mm wide, the terminal spike staminate, lateral ones androgynous or pistillate; bracts leaf-like, the lower 1–3 slightly longer than the inflorescence, long–sheathing at base; pistillate glumes brown, membranous, ovate, 2.5–5 mm long, margins hyaline, the midnerve prolonged into a scabrous awn 3–5 mm long; perigynium chartaceous to subcoriaceous, green to yellowish green, or reddish brown to dark red, ellipsoid to obovoid or broadly obovoid, (3–)5–8 mm long, 1–4 mm wide, many–veined, obtusely trigonous in cross section, the apex contracted gradually or abruptly to the beak, the apical teeth sharp, 0.5–3 mm long, base subcuneate to obtuse. Flowers unisexual. Corolla (petals) absent. Ovary superior; style 1–2 mm long, the base continuous to the achene apex without any thickening; stigmas 3.

Fruit: Achenes rhombic–ellipsoid; trigonous; without conspicuous constrictions at angles. Seeds 1; seed coat thin; free from pericarp; embryo basal; endosperm abundant.

Ploidy:

Habitat: Known from a single collection from Moanalua Valley.

Elevation Range: (10–)450–1,000 m.
360–370 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Other References

Wagner et al. 1990:1392 (K, O, Mo, L, M, H [as C. w. subsp. wahuensis], O [as C. w. subsp. herbstii]); Thomas, Magnacca, et al. 2025:239 (Syn. C. w. subsp. herbstii = C. w. subsp. wahuensis)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Carex wahuensis subsp. wahuensis Waianae Mts., Mt. Kaala, E. Slope, along trail Preserved_Specimen Krauss, R.W. 198 Oahu BISH 1948-04-18
2 Carex wahuensis subsp. wahuensis Waiakeakua Preserved_Specimen Munro, G.C. 157 Lanai BISH 1927-07-18
3 Carex wahuensis subsp. wahuensis N Pohakukane, Haena Preserved_Specimen Degener, O. 17648 Kauai BISH 1939-12-23
4 Carex wahuensis subsp. wahuensis Kaala Preserved_Specimen Degener, O. 8281 Oahu BISH 1928-02-11
5 Carex wahuensis subsp. wahuensis Ridge between Niu and Wailupe Preserved_Specimen Forbes, C.N. 2456 Oahu BISH 1917-04-11
6 Carex wahuensis subsp. wahuensis Kalalau trail, 1st 1-2 mi. Preserved_Specimen Wagner, W.L. 5180 Kauai BISH 1983-12-21
7 Carex wahuensis subsp. wahuensis Lanai Preserved_Specimen Munro, G.C. 171 Lanai BISH
8 Carex wahuensis subsp. wahuensis Waiakeakua Preserved_Specimen Munro, G.C. 157 Lanai BISH 1927-07-18
9 Carex wahuensis subsp. wahuensis Pupukea Trail Preserved_Specimen Lyon, H.L. Oahu BISH 1934-04-20
10 Carex wahuensis subsp. wahuensis N of Kaala Preserved_Specimen Degener, O. 17448 Oahu BISH 1937-04-26
11 Carex wahuensis subsp. wahuensis Mauka of McGregor Preserved_Specimen Degener, O. 22080 Maui BISH 1952-03-09
12 Carex wahuensis subsp. wahuensis Olowalu Valley Preserved_Specimen Forbes, C.N. 2339 Maui BISH 1920-05-12
13 Carex wahuensis subsp. wahuensis 3rd gulch of Hauula mountain range Preserved_Specimen Herat, T. 234 Oahu BISH 1972-08-24
14 Carex wahuensis subsp. wahuensis Koolau Mts., Pia Valley, ~2 mi into gulch Preserved_Specimen Sohmer, S.H. 6473 Oahu BISH 1970-06-10
15 Carex wahuensis subsp. wahuensis Kokee, Kilohana Rd. from Kalalau Lookout Preserved_Specimen Stone, B.C. 3410 Kauai BISH 1960-04-14
16 Carex wahuensis subsp. wahuensis Pohaku Preserved_Specimen Munro, G.C. 408 Lanai BISH 1915-02-27
17 Carex wahuensis subsp. wahuensis Waianae Mts., ridge to Palikea, 1/4 mi. along 1st main E ridge Preserved_Specimen Krauss, R.W. 161 Oahu BISH 1948-03-26
18 Carex wahuensis subsp. wahuensis Kaapahu, Old Donkey Trail on the Kalepa side under some lamas before Alalele Stream Preserved_Specimen Welton, P. 2334 Maui BISH 2013-01-10
19 Carex wahuensis subsp. wahuensis Waianae Mts., Puu Kanehoa, ridge to Preserved_Specimen Beardsley, J.W. Oahu BISH 1959-03-26
20 Carex wahuensis subsp. wahuensis Lanaihale Preserved_Specimen Kondo, Y. Lanai BISH 1961-03-25
21 Carex wahuensis subsp. wahuensis Palikea, Waianae Mts. Preserved_Specimen Pearsall, G.A. Oahu BISH 1954-07-08
22 Carex wahuensis subsp. wahuensis near power poles on crest between Kuliouou and Hahaione Valleys Preserved_Specimen Koes, E. Oahu BISH 1997-02-26
23 Carex wahuensis subsp. wahuensis Kipahulu Valley, between upper pasture and gauging station on Palakea Stream Preserved_Specimen Gagné, B.H. Maui BISH 1975-03-01
24 Carex wahuensis subsp. wahuensis Kaua Preserved_Specimen UH student Oahu BISH 1958-04-26
25 Carex wahuensis subsp. wahuensis Kokee Preserved_Specimen Smith, F.G. Kauai BISH 1929-05-01
26 Carex wahuensis subsp. wahuensis Oio Paumalu trail Preserved_Specimen Degener, O. 17646 Oahu BISH 1940-06-16
27 Carex wahuensis subsp. wahuensis Pupukea Preserved_Specimen Meebold, A. Oahu BISH 1932-05-01
28 Carex wahuensis subsp. wahuensis Preserved_Specimen Rémy, J. Hawaii BISH 1851-01-01
29 Carex wahuensis subsp. wahuensis Olowalu Valley Preserved_Specimen Forbes, C.N. 2338 Maui BISH 1920-05-12
30 Carex wahuensis subsp. wahuensis Preserved_Specimen Degener, O. 8368 BISH
31 Carex wahuensis subsp. wahuensis Preserved_Specimen Degener, O. 8369 BISH
32 Carex wahuensis subsp. wahuensis Preserved_Specimen Degener, O. 8367 BISH
33 Carex wahuensis subsp. wahuensis Waianae Mts., central Lualualei, below Kanehoa, valley 4 slopes of gully Preserved_Specimen herb Christophersen, E. 3677 Oahu BISH 1932-06-01
34 Carex wahuensis subsp. wahuensis Olowalu Valley Preserved_Specimen Forbes, C.N. 2285 Maui BISH 1920-05-09
35 Carex wahuensis subsp. wahuensis Pohakuloa Preserved_Specimen Munro, G.C. 257 Lanai BISH 1915-02-27
36 Carex wahuensis subsp. wahuensis Lanihuli trail Preserved_Specimen Forbes, C.N. 1471 Oahu BISH 1910-03-15
37 Carex wahuensis subsp. wahuensis Above Nuuanu Pali Preserved_Specimen Degener, O. 8277 Oahu BISH 1926-10-31
38 Carex wahuensis subsp. wahuensis Wailupe Preserved_Specimen Forbes, C.N. 2523 Oahu BISH 1919-01-01
39 Carex wahuensis subsp. wahuensis Waianae Mts., trail to Palikea Preserved_Specimen Krauss, R.W. 108 Oahu BISH 1948-02-04
40 Carex wahuensis subsp. wahuensis Waianae Mts., to Palikea, forest trail No. 19 With grasses and Styphelia -- open ridge line Preserved_Specimen Krauss, R.W. 109 Oahu BISH 1948-02-04
41 Carex wahuensis subsp. wahuensis Kaena Point, top dry, rocks Preserved_Specimen Degener, O. 8220 Oahu BISH 1932-01-31
42 Carex wahuensis subsp. wahuensis Hale Pohaku, Mauna Kea, Hamakua District Growing in rocky substrate with Deschampsia australis, Stipa cernua, and Trisetum glomeratum in open mamane parkland. Preserved_Specimen Sedge about 1 ft tall Cuddihy, L.W. 897 Hawaii BISH 1981-09-06
43 Carex wahuensis subsp. wahuensis Waianae Mts., to Palikea, forest trail No. 19 Forest trail Preserved_Specimen Krauss, R.W. 111 Oahu BISH 1948-02-04
44 Carex wahuensis subsp. wahuensis W Maui, slopes of Helu Peak Gowing alongside typical shiny light-green leaved plants. Preserved_Specimen Robust glaucus -blue leaved plants. Hobdy, R.W. 4955 Maui BISH 2006-01-27
45 Carex wahuensis subsp. wahuensis Makaha, gulch W of Puu Keaau low thicket of Dodonaea and Bidens waianensis Preserved_Specimen St.John, H. 25508 Oahu BISH 1955-01-29
46 Carex wahuensis subsp. wahuensis Waimea District. Waimea Canyon, just southeast of Kukui peak. Acacia-Metrosideros lowland dry to mesic forest with Dodonaea viscosa, Leptecophylla tameiameiae, Hibiscus waimeae, Diospyros sandwicensis, Chrysodracon aurea (=C. stenophylla), Planchonella sandwicensis, Nestegis sandwicensis, Bobea brevipes, Sida Preserved_Specimen Sedge, terrestrial, leaves glossy dark green, one of two forms of C. wahuensis subsp. Wahuensis in immediate area, the other form (Wood 16352) is larger with green-blue leaves, both are common. Wood, K.R. 16349 Kauai BISH 2015-03-05
47 Carex wahuensis subsp. wahuensis Kipapa Gulch, S ridge, Koolau Mts., Waipio moist woods Preserved_Specimen Hosaka, E.Y. 948 Oahu BISH 1933-03-19
48 Carex wahuensis subsp. wahuensis S of Maunahui cabin sunny ravine Preserved_Specimen Degener, O. 8230 Molokai BISH 1928-04-15
49 Carex wahuensis subsp. wahuensis NE of Kipu woods Preserved_Specimen Degener, O. 2205 Kauai BISH 1926-06-17
50 Carex wahuensis subsp. wahuensis Waianae Mts., Honouliuli, Puu Kanehoa wooded ridge Preserved_Specimen tufts St.John, H. 11089 Oahu BISH 1931-03-22
51 Carex wahuensis subsp. wahuensis East Maui, Hana District, Haleakala, Kaupo trail eroded lava flows Preserved_Specimen 24 in. Olson, G.E. 55 Maui BISH 1937-07-25
52 Carex wahuensis subsp. wahuensis Ridge E of Central Makeleha Valley dry, grassy; with Dodonaea, Bidens, Diospyros Preserved_Specimen Ozaki, E.T. 1508 Oahu BISH 1956-04-15
53 Carex wahuensis subsp. wahuensis Koolau Mts, Moanalua valley, first ridge makai of main valley divide On cliff with Eragrostis variabilis, Carex meyenii, Metrosideros, Sida fallax, Chamaesyce, Psilotum nudum. Preserved_Specimen Perlman, S. 14088 Oahu BISH 1994-03-22
54 Carex wahuensis subsp. wahuensis Manoa Valley, ridge above Woodlawn dry, Lantana covered slope Preserved_Specimen Fosberg, F.R. 9296 Oahu BISH 1933-03-19
55 Carex wahuensis subsp. wahuensis Waimea District. Kumuwela, slopes west of Poomau stream Metrosideros Cheirodendron montane wet forest, 80-95% closed, 8-12 m tall canopy, some emergent Metrosieros 20 m tall Psychotria mariniana, Santalum pyruliarium, Bobea brevipes, Kadua affinis, Coprosma waimeae, Melicope barbigera, Dodonaea viscosa, Dianella sandwicensis, Sphenomeris chinensis, Doodia kunthiana, Dropt Preserved_Specimen sedge, teresstiral Wood, K.R. 15342 Kauai BISH 2013-01-10
56 Carex wahuensis subsp. wahuensis Waianae Mts., trail to Puu Kaua dry, NE facing slope Preserved_Specimen Krauss, R.W. 121 Oahu BISH 1948-02-29
57 Carex wahuensis subsp. wahuensis Koolau Range, Palolo Valley koa forest, steep slopes Preserved_Specimen Fosberg, F.R. 9273 Oahu BISH 1933-03-19
58 Carex wahuensis subsp. wahuensis Pali grassy slope Preserved_Specimen Hitchcock, A.S. 13788 Oahu BISH 1916-06-17
59 Carex wahuensis subsp. wahuensis Waianae Mts., NW of Puu Kanehoa edge of forest Preserved_Specimen tufts St.John, H. 14018 Oahu BISH 1934-01-07
60 Carex wahuensis subsp. wahuensis West Maui, Lahaina District, Lihau Natural Area Reserve, SE side of planze near Olowalu Valley Occasional, with common wider-leaved form, on sunny, shrubby ridgetop Preserved_Specimen Oppenheimer, H.L. 31305 Maui BISH 2013-03-13
61 Carex wahuensis subsp. wahuensis S of Waihee dry forest in forehills Preserved_Specimen Degener, O. 21960 Maui BISH 1952-02-28
62 Carex wahuensis subsp. wahuensis Tantalus exposed ridge Preserved_Specimen Degener, O. 8274 Oahu BISH 1926-03-28
63 Carex wahuensis subsp. wahuensis Koolau Mts, Moanalua valley, back of valley, first ridge makai of main divede, on small knoll In Metrosideros-Diospyros lowland mesic forest and shrubland with Phyllanthus, Chamaesyce, Eragrostis variabilis, Sida, Peperomia, Osteomeles, Andropogon, Erigeron karvinskianus, Ageratina riparia, Canthium. Preserved_Specimen Plants with flowers and fruits. Perlman, S. 12979 Oahu BISH 1992-09-23
64 Carex wahuensis subsp. wahuensis Kolekole Pass exposed ridge Preserved_Specimen Degener, O. 8267 Oahu BISH 1927-01-01
65 Carex wahuensis subsp. wahuensis west Maui, Lahaina District, Ukumehame, west fork of Papalaua Gulch, near Hanaula Iki. Vegetation: Metrosideros, Acacia, Planchonella, Nestegis, Leptecophylla, Chrysodracon, Dodonaea, Euphorbia, Myoporum, Santalum, Coprosma, Xylosma, Psychotria, Alyxia, Dryopteris, Panicum, Eragrostis, Carex, Doodia, Bidens, Peperomia, Lantana, Erigeron, Argeratina, Oplismenus. Preserved_Specimen Locally common glaucous form, more common than typical form with glossy, dark green leaves. Oppenheimer, H.L. 41604 Maui BISH 2016-04-08
66 Carex wahuensis subsp. wahuensis Waianae Mts., gulch above Puu Hapapa, ca. 100 yds above contour trail. Wet, organic soil on bottom of gulch. Somewhat open area. Preserved_Specimen ferns, mosses, banana. Herbst, D.R. 66 Oahu BISH 1966-04-23
67 Carex wahuensis subsp. wahuensis East Maui, Haleakala, Kaupo Gap, 1st cove S of Waikekeehia grass on dry basalt gravel Preserved_Specimen tufts St.John, H. 21207 Maui BISH 1945-08-27
68 Carex wahuensis subsp. wahuensis Waianae Mts., 1st gully below Peacock Flats dry; with Acacia koa, Psidium, Aleurites, Sapindus oahuensis, Osmanthus sandwicensis, Eugenia cumini, Schinus terebinthifolius Preserved_Specimen Herbst, D.R. 322 Oahu BISH 1967-02-25
69 Carex wahuensis subsp. wahuensis Wailau Valley Pandanus forest, moist gully Preserved_Specimen Fosberg, F.R. 13084 Molokai BISH 1936-04-19
70 Carex wahuensis subsp. wahuensis W Maui, Mahinahina Gulch Mesic forest Preserved_Specimen Perennial to 80cm Annable, C.R. 3933 Maui BISH 1999-02-19
71 Carex wahuensis subsp. wahuensis Outside forest reserve, along footpath trail leading to Sacred Falls Along footpath. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
72 Carex wahuensis subsp. wahuensis Awaawapuhi trail, Honopu moist Acacia forest Preserved_Specimen St.John, H. 26015 Kauai BISH 1956-12-29
73 Carex wahuensis subsp. wahuensis Bird Park (Kipuka Puaulu) moderate forest Preserved_Specimen Morley, H. 155 Hawaii BISH 1934-08-21
74 Carex wahuensis subsp. wahuensis Waimea, Paaiki Valley, N of Milolii Ridge Rd. dry, shady floor of canyon; with Isodendrion subsessilifolium, Pisonia sandwicensis, Aleurites molucanna, Antidesma platyphyllum, Lantana Preserved_Specimen Gustafson, R. 1751 Kauai BISH 1980-05-25
75 Carex wahuensis subsp. wahuensis Awaawapuhi, along trail Metrosideros polymorpha montane mesic forest. W/Psychotria mariniana, Melicope anisata, M. barbigera, Pouteria sandwicensis, dodonaea viscosa, Dianella sandwicensis, Tetraplasandra kavaiensis, Claoxylon sandwicensis Preserved_Specimen terrestrial sedge, mature perigynium yellow-brown, leaves with green-blue tinge, common Wood, K.R. 14694 Kauai BISH 2011-06-29
76 Carex wahuensis subsp. wahuensis Munro Trail, on rainy top among bushes on rainy top Preserved_Specimen Degener, O. 24303 Lanai BISH 1957-08-13
77 Carex wahuensis subsp. wahuensis Napali coast, E of Hoolulu Valley base of cliffs, dry forest Preserved_Specimen Christensen, C. 176 Kauai BISH 1977-03-30
78 Carex wahuensis subsp. wahuensis Kipapa gulch wet forest Preserved_Specimen Fosberg, F.R. 9790 Oahu BISH 1933-08-08
79 Carex wahuensis subsp. wahuensis Waimea, Milolii Ridge open scrub between scattered Acacia koa Preserved_Specimen St.John, H. 13717 Kauai BISH 1933-12-27
80 Carex wahuensis subsp. wahuensis Mt. Kaala, DuPont trail open Acacia forest Preserved_Specimen tufted St.John, H. 23689 Oahu BISH 1949-05-22
81 Carex wahuensis subsp. wahuensis trail from Paliku to NE rim of Haleakala dry slope Preserved_Specimen Degener, O. 12555 Maui BISH 1939-08-07
82 Carex wahuensis subsp. wahuensis Kalulu, Kapano Gulch woods Preserved_Specimen tufted St.John, H. 18764 Lanai BISH 1938-04-10
83 Carex wahuensis subsp. wahuensis Na Pali Coast, Hanakoa Valley Growing along trail Preserved_Specimen Sedge 1.5-2 ft tall Corn, C. 184 Kauai BISH 1980-04-10
84 Carex wahuensis subsp. wahuensis Waianae Mts., Honouliuli open ridge Preserved_Specimen tufts St.John, H. 11044 Oahu BISH 1931-03-22
85 Carex wahuensis subsp. wahuensis Waianae Mts., to Palikea, forest trail No. 19 Forest trail Preserved_Specimen Krauss, R.W. 114 Oahu BISH 1948-02-04
86 Carex wahuensis subsp. wahuensis Nuuanu Pali Growing sympatrically with Carex wahuensis subsp. wahuensis (see Perlman 21103). Preserved_Specimen Perlman, S. 21101 Oahu BISH 2008-08-06
87 Carex wahuensis subsp. wahuensis Koolau Mts., Laie trail, Kahawainui Gulch forest Preserved_Specimen large sedge Stone, B.C. 1234 Oahu BISH 1956-03-25
88 Carex wahuensis subsp. wahuensis Waialua District. Waianae Mountains, Pupukea-Paumalu Forest Reserve, up from Boy Scout Camp. Degraded Psidium cattleianum dominant forest W/Psidium cattleianum, big dead downed Acacia koa, Grevillea, Carex, Dianella, Clidemia, Alyxia, Dicranopteris. Preserved_Specimen New leaves light green, older dark green. Edges and midrib of underside of leaf rough. Clumping. Inflo upright. Tangalin, N. 3145 Oahu BISH 2012-04-22
89 Carex wahuensis subsp. wahuensis Kolekole Pass dry Preserved_Specimen Degener, O. 5268 Oahu BISH 1926-05-01
90 Carex wahuensis subsp. wahuensis Waianae Mts., Mt. Kaala, E slope trail Preserved_Specimen Krauss, R.W. 199 Oahu BISH 1948-04-18
91 Carex wahuensis subsp. wahuensis Hanalei, Kalalau, Lower valley, up Southwestern drainage below Kalahu Psydrax degraded mesic forest dominated by invasive Syzygium cumini, w/ cliff species of Lobelia niihauensis, Doryopteris decipiens, Carex wahuensis, Carex meyenii, Wikstroemia oahuensis, Lipochaeta connata acris, Artemisia australis, Euphorbia celastroides hanapepen... Preserved_Specimen Sedge, occasional on cliffs w/ C. Meyenii, w/ fruit. K.R. Wood 19498 Kauai BISH 2024-05-21
92 Carex wahuensis subsp. wahuensis Koolau Range, above Pupukea trail, summit trail, ~200 yds beyond shelter edge of grassy clearing Preserved_Specimen Pearsall, G.A. 386 Oahu BISH 1959-05-04
93 Carex wahuensis subsp. wahuensis Olokele ditch trail, Makaweli, 2 mi below ditch house dry open basalt banks Preserved_Specimen tufts St.John, H. 23087 Kauai BISH 1947-12-26
94 Carex wahuensis subsp. wahuensis Wailuku, Iao Valley moist gorge Preserved_Specimen St.John, H. 10280 Maui BISH 1930-02-09
95 Carex wahuensis subsp. wahuensis Olinda pipe line trail open, wet rain forest Preserved_Specimen Degener, O. 8249 Maui BISH 1927-06-22
96 Carex wahuensis subsp. wahuensis Waianae Mts., Puu Kalena, Waianaeuka Metrosideros forest Preserved_Specimen tufted St.John, H. 22564 Oahu BISH 1947-02-16
97 Carex wahuensis subsp. wahuensis Kipapa Trail, ~2/3 mi from refuge boundary, N side of trail disturbed wet lowland forest; Metrosideros, Acacia, Dicranopteris, Psidium cattleianum Preserved_Specimen ~75 cm sedge; inflorescence spike-like; perigynium yellow Reynolds, L.S. 138 Oahu BISH 2006-06-13
98 Carex wahuensis subsp. wahuensis Waianae Mts., Puu Hapapa, Honouliuli wooded valley Preserved_Specimen tufts St.John, H. 10441 Oahu BISH 1930-03-16
99 Carex wahuensis subsp. wahuensis Tantalus rain forest Preserved_Specimen Degener, O. 8275 Oahu BISH 1926-03-20
100 Carex wahuensis subsp. wahuensis Waiakaiole drainage (headwaters of Waiapaa Gulch), edge of shallow stream bank on 15 degree slope, N aspect Lowland Dry Shrubland/Forest. Growing on edge of shallow, red clay stream bank on 15 degree slope, N aspect, with dappled sunlight 50% of day, mesic conditions. Associated with Syzygium jambos, Psidium cattleianum, Blechnum appendiculatum, Microlepia strigosa, Leucaena leucocephala, Solanum americanum, Conyza bonariensis, Passiflora suberosa, Sadleria cyatheoides, Dianella sandwicensis. Preserved_Specimen Rhizomatous, clumping sedge. In flower and fruit; terminal flower spikes staminate, lower pistillate. Normal vigor. Rare in area. 2 mature, ca 100 seedlings in 5m x 5m area. LeGrande, M. 1274 Lanai BISH 2003-02-06
101 Carex wahuensis subsp. wahuensis Napali trail between Makana and Waiahuakua grassy slope above ocean Preserved_Specimen Degener, O. 17585 Kauai BISH 1939-12-24
102 Carex wahuensis subsp. wahuensis Koaie Canyon, rocky ledge near base of S facing wall protected by roots of Metrosideros polymorpha Preserved_Specimen sedge, blades to 80 cm Trauernicht, C. 722 Kauai BISH 2009-02-02
103 Carex wahuensis subsp. wahuensis Mauumae ridge with Clidemia, Dicranopteris, Psychotria Preserved_Specimen clump Takeuchi, W.N. 102 Oahu BISH 1984-03-10
104 Carex wahuensis subsp. wahuensis Waianae Mts., trail to Mt. Kaala In shade Preserved_Specimen Single plant; frt. bright yellow. Krauss, R.W. 183 Oahu BISH 1948-04-18
105 Carex wahuensis subsp. wahuensis 1st gully makai S side of Keaau Valley dry Preserved_Specimen Degener, O. 8219 Oahu BISH 1932-02-07
106 Carex wahuensis subsp. wahuensis Site 22 OS, Piihonua, Hilo Forest Reserve Open healthy. Associated spp.: Metrosideros polymorpha, Ilex anomala, Gouldia sp., Pelea clusiifolia, Cheirodendron trigynum, Myrsine lessertiana, Freycinetia arborea, Alyxia oliviformis. Soils are deep and moderately well drained ash, substrate unknown. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-19
107 Carex wahuensis subsp. wahuensis Waianae Mts., ridge trail to Palikea dry Preserved_Specimen Cowan, R.S. 814 Oahu BISH 1948-02-04
108 Carex wahuensis subsp. wahuensis Pupukea-Kahuku trail, Pupukea side open ridge Preserved_Specimen Degener, O. 8221 Oahu BISH 1929-03-30
109 Carex wahuensis subsp. wahuensis Waianae Mts., Puu Kaua grassy edge of cliff Preserved_Specimen tufted St.John, H. 20022 Oahu BISH 1939-03-05
110 Carex wahuensis subsp. wahuensis Mauna Kea, near Puu Laau Cabin, Hamakua District Growing with Holcus lanatus and Dactylis glomerata in open mamane parkland. Preserved_Specimen Sedge 1-1.5 ft tall Cuddihy, L.W. 869 Hawaii BISH 1981-09-24
111 Carex wahuensis subsp. wahuensis Kokee Camp moist ravine Preserved_Specimen Degener, O. 2211 Kauai BISH 1926-06-23
112 Carex wahuensis subsp. wahuensis W Maui, N of Pohakea Gulch ridge; rain forest Preserved_Specimen Degener, O. 8247 Maui BISH 1927-07-23
113 Carex wahuensis subsp. wahuensis Koolau Range, Ewa FR, Kalauao Ridge wooded ridge Preserved_Specimen tufts St.John, H. 13037 Oahu BISH 1933-03-29
114 Carex wahuensis subsp. wahuensis West Maui, Maunalei Arboretum on Mahana Ridge Trail to Puu kaeo lookout. Disturbed Mesic Metrosideros-Ardisia Forest: with Metrosideros polymorpha, M. waialealae, Dicranopteris, Nephrolepis, Ardisia, Cyanea angustifolia, Clermontia kakeana, Wikstroemia oahuensis, Cinnamomum, Psilotum nudum. Preserved_Specimen Occasional on forest floor with spider living in leaf hunting Hyposcocoma. Green immature seed and white stamens on fruiting head. Tangalin, N. 3490 Maui BISH 2013-04-12
115 Carex wahuensis subsp. wahuensis Southwest flank of Mauna Loa, in the ahupuaa of Manuka Vegetation: Semi Closed Metrosideros forest on soil and weathered aa. Understory consisting of Cibotium, Sadleria, Astelia and Myrsine. Preserved_Specimen Higashino, P.K. 6276 Hawaii BISH 1977-07-22
116 Carex wahuensis subsp. wahuensis Mahana, Hulopoe Gulch moist thicket Preserved_Specimen tufts St.John, H. 18736 Lanai BISH 1938-04-09
117 Carex wahuensis subsp. wahuensis Palikea, Honolulu side small valley Preserved_Specimen Degener, O. 8218 Oahu BISH 1932-10-23
118 Carex wahuensis subsp. wahuensis East Maui, Makawao District, Haleakala, near jct. Halemauu trail and park road scrub, small open ridge Preserved_Specimen Henrickson, J.S. 3852 Maui BISH 1969-07-13
119 Carex wahuensis subsp. wahuensis Koolau Range, Wiliwilinui ridge under Psidium thicket Preserved_Specimen Krauss, R.W. 247 Oahu BISH 1948-06-28
120 Carex wahuensis subsp. wahuensis Hanalei District. coastal cliffs below Princeville, Kenomene region secondary invasive vegetation, with some relic native taxa native relics include Pandanus tectorius, Scaevola taccada, Vigna marina, Bacopa monnieri, schoenoplectus tabernaemontani, Cyperus javanicus, Fimbristylis cymosa subsp. spathacea, Portulaca lutea, Lys Preserved_Specimen sedge, terrestrial on steep rocky slope above sea, immature perigynium green, maturing yellow, scattered locations but uncommon Wood, K.R. 15533 Kauai BISH 2013-06-20
121 Carex wahuensis subsp. wahuensis Waianae Mts., Mt. Kaala, E. slope, along trail trail, E. slope Preserved_Specimen Fls. yellow Krauss, R.W. 197 Oahu BISH 1948-04-18
122 Carex wahuensis subsp. wahuensis Kaohai, Awehi Gulch, head of woods Preserved_Specimen St.John, H. 18888 Lanai BISH 1938-04-15
123 Carex wahuensis subsp. wahuensis Haleakala, Puu Oili moist old lava flows Preserved_Specimen 18 in. Olson, G.E. 15 Maui BISH 1936-12-01
124 Carex wahuensis subsp. wahuensis below Puu Kapu dry ridge Preserved_Specimen Christophersen, E. 1549 Oahu BISH 1931-02-12
125 Carex wahuensis subsp. wahuensis Kipapa Gulch, S ridge, Koolau Mts., Waipio open ridge, moist Preserved_Specimen Hosaka, E.Y. 540 Oahu BISH 1932-05-15
126 Carex wahuensis subsp. wahuensis Mahanaloa Valley, above Kuia Valley, ~2.5 mi W of Hwy 55 valley bottom with kukui, Diospyros, Pelea, Pleomele Preserved_Specimen tufted, tussock forming; inflorescence greenish Lorence, D.H. 5112 Kauai BISH 1967-03-10
127 Carex wahuensis subsp. wahuensis NW of Maunahui small ravine; dry, shady embankment Preserved_Specimen Degener, O. 8239 Molokai BISH 1928-05-20
128 Carex wahuensis subsp. wahuensis Waianae Mts., Mt. Kaala, E slope trail, shade Preserved_Specimen Krauss, R.W. 188 Oahu BISH 1948-04-18
129 Carex wahuensis subsp. wahuensis S Kahanahaiki Valley dry slope Preserved_Specimen Degener, O. 17433 Oahu BISH 1931-11-01
130 Carex wahuensis subsp. wahuensis Wailau Valley talus at foot of cliff Preserved_Specimen Fosberg, F.R. 9679 Molokai BISH 1933-07-04
131 Carex wahuensis subsp. wahuensis Pahole Ridge; between drainage #4 & #5 Under Acacia koa. Preserved_Specimen Welton, P. 500 Oahu BISH 1990-06-05
132 Carex wahuensis subsp. wahuensis W Maui, Puu Anu, Manawainui Gulch dry Preserved_Specimen Degener, O. 8243 Maui BISH 1927-07-12
133 Carex wahuensis subsp. wahuensis Waianae Mts. Makaha-Waianae Kai-Puu Kawiwi-Kaala ridge wooded Preserved_Specimen tufted St.John, H. 17612 Oahu BISH 1935-03-21
134 Carex wahuensis subsp. wahuensis Koolau Range, Hauula, Papali ridge from Cooper's Ranch koa forest Preserved_Specimen Krauss, R.W. 140 Oahu BISH 1948-03-21
135 Carex wahuensis subsp. wahuensis W Maui, N of Pohakea Gulch ridge, rain forest Preserved_Specimen Degener, O. 8246 Maui BISH 1927-07-23
136 Carex wahuensis subsp. wahuensis Upper Onini Gulch Occasional along edge of dry stream bed. Also present are Viola tracheliifolia, Doodia kunthiana, Dissochondrus biflorus, Selaginella, Pneumatopteris, Cyrtandra and Lysimachia. Preserved_Specimen Davis, J. 867 Molokai BISH 1982-05-28
137 Carex wahuensis subsp. wahuensis Mt. Haupu, on Kipu side, N side of Naluakeina With Lepidium orbiculare, Chamaesyce celastroides var. hanapepeensis, Heteropogon contortus, Metrosideros polymorpha, Bidens, Eragrostis variabilis, Peperomia, Lantana camara, Schinus terebinthifolius, Artemisia australis Preserved_Specimen Wood, K.R. 2815 Kauai BISH 1993-10-25
138 Carex wahuensis subsp. wahuensis East Maui, Haleakala, above Holua rock cliffs Preserved_Specimen Krauss, R.W. 348 Maui BISH 1948-09-03
139 Carex wahuensis subsp. wahuensis Koolau Range, Halawa Ridge Trail dry Preserved_Specimen Cowan, R.S. 955 Oahu BISH 1948-04-25
140 Carex wahuensis subsp. wahuensis Puhimau Crater Occasional on black cinder and ash soil in somewhat open Metrosideros collina forest with Gleichenia Preserved_Specimen Tufts, fruit yellow Fosberg, F.R. 56727 Hawaii BISH 1977-03-20
141 Carex wahuensis subsp. wahuensis Kahuaawi Gulch embankment Preserved_Specimen Degener, O. 8238 Molokai BISH 1928-05-12
142 Carex wahuensis subsp. wahuensis Olinda pipe line gulch, moist side Preserved_Specimen 0.75-1 m Lee, H.A. 184 Maui BISH 1926-03-01
143 Carex wahuensis subsp. wahuensis Koolau Range, Laie-Malaekahana ridge woods Preserved_Specimen tufts St.John, H. 13047 Oahu BISH 1933-04-15
144 Carex wahuensis subsp. wahuensis Waianae Mts., Honouliuli FR, Pohakea Pass grassy slope Preserved_Specimen tufts St.John, H. 13154 Oahu BISH 1933-05-12
145 Carex wahuensis subsp. wahuensis W Maui, slopes of Helu Peak In native shrubland. Preserved_Specimen Plants bright green with shiny green leaves. Hobdy, R.W. 4954 Maui BISH 2006-01-27
146 Carex wahuensis subsp. wahuensis W Maui, Puu Anu; Manawainui Gulch dryish region. Preserved_Specimen Degener, O. 8244 Maui BISH 1927-07-12
147 Carex wahuensis subsp. wahuensis East Maui, Hana District, Waihoi Valley stream bank, above Joinvillea stand Preserved_Specimen Harrison, B. 2583 Maui BISH 1972-01-01
148 Carex wahuensis subsp. wahuensis Lualualei open slopes, bottom of gully; under Bobea sandwicensis Preserved_Specimen Takeuchi, W.N. 3276 Oahu BISH 1987-01-18
149 Carex wahuensis subsp. wahuensis East Maui, Haleakala, Koolau Gap moist wall Preserved_Specimen Wilbur, R.L. 951 Maui BISH 1948-07-02
150 Carex wahuensis subsp. wahuensis Kipu, Haupu base of cliff, grassy cleft Preserved_Specimen tufts St.John, H. 13619 Kauai BISH 1933-12-25
151 Carex wahuensis subsp. wahuensis Kokee State Park, Kauai, Hawaii Growing in dense forest, mesic to wet conditions. Preserved_Specimen Compact, clumping sedge. Crago, L.M. 2005163 Kauai BISH 2005-08-03
152 Carex wahuensis subsp. wahuensis Napali Coast, Haena-Kalalau trail, Waiahuakua valley shady, forested, near stream Preserved_Specimen sedge Stone, B.C. 839 Kauai BISH 1955-06-18
153 Carex wahuensis subsp. wahuensis Waimea District, Kohua Ridge North facing slopes above Awini Stream Metrosideros-Acacia montane mesic forest with Alphitonia ponderosa, Antidesma platyphylla, Euphorbia atrococca, Claoxylon sandwicensis, Dodonaea viscosa, Myrsine lanaiensis, Nestegis sandwicensis, Dracaena aurea, Polyscias Kavaiensis, Planch Preserved_Specimen sedge, terrestial, common Wood, K.R. 14986 Kauai BISH 2012-05-17
154 Carex wahuensis subsp. wahuensis Waianae Mts., Honouliuli, N fork of valley E of Palikea moist shaded rock Preserved_Specimen tuft St.John, H. 10385 Oahu BISH 1930-02-23
155 Carex wahuensis subsp. wahuensis Kaumokunui Gulch, Mokileia Gulch. Preserved_Specimen Degener, O. 17447 Oahu BISH 1936-04-13
156 Carex wahuensis subsp. wahuensis Kamokou Preserve, ridge between Kupai`a and Kaunakakai Gulches, East Moloka`i. Dry shrubland of native shrubs and both native and alien grasses. Preserved_Specimen Sedge with red-brown fruit. Pratt, L.W. 1242 Molokai BISH 1982-05-30
157 Carex wahuensis subsp. wahuensis Kamoku, Kaiholena Gulch, SW ridge moist woods Preserved_Specimen tufts St.John, H. 22609 Lanai BISH 1947-04-04
158 Carex wahuensis subsp. wahuensis W Maui, Lahaina District, Lihau Lowland Ohia Mixed Mesic Shrubland at 3000 ft with low statured Metrosideros, Dodonaea, Styphelia, Wikstroemia, Osteomeles, Dubautia linearis, Gahnia, Pityrogramma, Sphenomeris, Pteridium, Trisetum inaequale, Cocculus, Eragrostis grandis and Psilotum Preserved_Specimen Welton, P. 1503 Maui BISH 1992-01-10
159 Carex wahuensis subsp. wahuensis NW Alakai swamp, Napali-Kona FR woods Preserved_Specimen St.John, H. 10766 Kauai BISH 1930-12-27
160 Carex wahuensis subsp. wahuensis 6.75 miles e. of Puu Huluhulu, Saddle Road mossy Metrosideros forest in kipuka Preserved_Specimen St.John, H. 25337 Hawaii BISH 1954-12-25
161 Carex wahuensis subsp. wahuensis Hamakua Dist.; Kailikaula Cliffs and stream Metrosideros-Osteomeles-Chamaesyce lowland mesic forest and cliff w/ riparian communities. Growing in basalt talus. Associated species: Carex spp., Dianella, Canthium, Eragrostis, Diospyros, Dodonaea, Alyxia. Preserved_Specimen Perigynum green; culms yellow-white. Wood, K.R. 4870 Hawaii BISH 1995-12-14
162 Carex wahuensis subsp. wahuensis W Maui, Lihau Collected on a dry S facing ridge. Preserved_Specimen Welton, P. 1351 Maui BISH 1991-10-26
163 Carex wahuensis subsp. wahuensis Kapapala Forest Reserve Growing with Dryopteris, and Athyrium spp., in closed Acacia koa - Metrosideros forest. Preserved_Specimen Occasional herb. Davis, J. 557 Hawaii BISH 1981-07-01
164 Carex wahuensis subsp. wahuensis Waialae Nui ridge top, dry; with Acacia koa, Metrosideros polymorpha, Psidium cattleianum, Diospyros sandwicensis Preserved_Specimen Takeuchi, W.N. 467 Oahu BISH 1985-01-26
165 Carex wahuensis subsp. wahuensis Nualolo trail, Napali-Kona FR woods Preserved_Specimen St.John, H. 10817 Kauai BISH 1930-12-28
166 Carex wahuensis subsp. wahuensis Waimea District. Waimea Canyon, just southeast of Kukui peak. Acacia-Metrosideros lowland dry to mesic forest with Dodonaea viscosa, Leptecophylla tameiameiae, Hibiscus waimeae, Diospyros sandwicensis, Chrysodracon aurea (=C. stenophylla), Planchonella sandwicensis, Nestegis sandwicensis, Bobea brevipes, Sida Preserved_Specimen Sedge, terrestriqal, leaves green-blue, one of two forms of C. wahuensis subsp. Wahuensis in immediate area, the other form (Wood 16349) is smaller with glossy dark green leaves, both are common. Wood, K.R. 16352 Kauai BISH 2015-03-05
167 Carex wahuensis subsp. wahuensis W Maui, Lahaina District, Lihau Lowland Koa Mesic Forest at 3250 ft on a ridge beneath a of canopy of 7- 8m Acacia koa, Psychotria, Dodonaea, Coprosma and Myrsine with a understory of native ferns, sedges, Trisetum inaequale and Liparis Preserved_Specimen Welton, P. 935 Maui BISH 1991-07-09
168 Carex wahuensis subsp. wahuensis W Maui, Lahaina Distr, Kauaula Valley, S side Locally common on ridgetop, in area burned in February 2007. Veg: Metrosideros, Sophora, Dodonaea, Leptecophylla, Eragrostis, Panicum, Vaccinium, Sadleria, Dianella, Pteridium, Bidens, Morelotia, Cocculus, Osteomeles, Andropogon, Senecio, Conyza, Melinis. Preserved_Specimen Growing with typical form, but distinct with its glaucous lvs. Oppenheimer, H.L. 21010 Maui BISH 2010-02-16
169 Carex wahuensis subsp. wahuensis Waianae Mts., trail to Puu Kaua dry, NE facing slope Preserved_Specimen Krauss, R.W. 123 Oahu BISH 1948-02-29
170 Carex wahuensis subsp. wahuensis East Maui, Hanawi, E ridge from Frisbee Meadow Metrosideros-Cheirodendron montane wet forest with Melicope clusiifolia, Dryopteris spp., Stenogyne rotundifolia, Polysticum bonseyi, Rubus macraei, Rubus hawaiensis, Lycopodium venustulum. On basalt vertical walls. Threats: Holcus lanatus, Hypochoeris radicata, Prunella vulgaris. Preserved_Specimen Clumping. Wood, K.R. 4313 Maui BISH 1995-06-23
171 Carex wahuensis subsp. wahuensis Waianae Mts., Puu Kaua grassy main divide Preserved_Specimen tufted St.John, H. 24052 Oahu BISH 1950-02-26
172 Carex wahuensis subsp. wahuensis Ekahanui Gulch, E of Puu Kaua, Waianae Mts, Honouliuli steep, moist ridge Preserved_Specimen tufts; ripe fruit yellow Fosberg, F.R. 29642 Oahu BISH 1948-02-29
173 Carex wahuensis subsp. wahuensis Puu Kanehoa, Waianae range dry wooded ridge Preserved_Specimen Hume, E.P. 482 Oahu BISH 1932-01-18
174 Carex wahuensis subsp. wahuensis East Maui, Kipahulu Valley, lower dogleg with koa, Perrottetia sandwicensis, Psychotria hawaiiensis, Broussaisia arguta, Hedyotis acuminata Preserved_Specimen infructescence green Wagner, W.L. 5904 Maui BISH 1988-03-09
175 Carex wahuensis subsp. wahuensis Kaala moist forest Preserved_Specimen ~1 m erect Bryan Jr., E.H. 706 Oahu BISH 1929-04-21
176 Carex wahuensis subsp. wahuensis Waianae Mts., Mokuleia mauka forest Preserved_Specimen Degener, O. 19581 Oahu BISH 1949-05-02
177 Carex wahuensis subsp. wahuensis Waimanalo base of cliff, steep ridge, thicket Preserved_Specimen tuft St.John, H. 21569 Oahu BISH 1946-03-24
178 Carex wahuensis subsp. wahuensis SW of Haena, !/2 mi in on Napali Coast trail exposed, crevices, black lava Preserved_Specimen Henrickson, J.S. 3990 Kauai BISH 1969-07-31
179 Carex wahuensis subsp. wahuensis Waianae Mts., Mauna Kapu, head of Nanakuli Valley wooded cliff Preserved_Specimen small clumps Fosberg, F.R. 13799 Oahu BISH 1937-05-02
180 Carex wahuensis subsp. wahuensis First ridge makai of main divide on the north side of Moanalua Valley Growing in cracks on an exposed cliff face with Eragrostis, Bidens, Phyllanthus, and Psilotum. Preserved_Specimen Herbst, D.R. 1495 Oahu BISH 1970-01-11
181 Carex wahuensis subsp. wahuensis NE ridge, Olokui, Wailau dense forest, steep ridge Preserved_Specimen St.John, H. 23291 Molokai BISH 1948-02-06
182 Carex wahuensis subsp. wahuensis Kuliouou, Koolau Mts. forest Preserved_Specimen Christophersen, E. 1526 Oahu BISH 1931-02-05
183 Carex wahuensis subsp. wahuensis W of Keopaweo, Nawiliwili dry forest Preserved_Specimen Degener, O. 21536 Kauai BISH 1952-01-05
184 Carex wahuensis subsp. wahuensis Hamakua Dist.; Kailikaula Cliffs and stream Metrosideros-Osteomeles-Chamaesyce lowland mesic forest and cliff w/ riparian communities. Growing in basalt talus. Associated species: Carex spp., Dianella, Canthium, Eragrostis, Diospyros, Dodonaea, Alyxia. Preserved_Specimen Perigynum yellow. Wood, K.R. 4871 Hawaii BISH 1995-12-14
185 Carex wahuensis subsp. wahuensis Waianae Mts., Waianaeuka, E ridge of Puu Kalena exposed Preserved_Specimen Fosberg, F.R. 13007 Oahu BISH 1936-03-22
186 Carex wahuensis subsp. wahuensis Top of Nuuanu Pali Road grassy slopes Preserved_Specimen clumps; staminate spikes brown, pistillate green Fosberg, F.R. 12281 Oahu BISH 1935-10-14
187 Carex wahuensis subsp. wahuensis E Manoa Valley dry woods Preserved_Specimen Degener, O. 8263 Oahu BISH 1927-02-06
188 Carex wahuensis subsp. wahuensis Kokee, Kumuwela lookout weeds Preserved_Specimen Stone, B.C. 3377 Kauai BISH 1960-04-12
189 Carex wahuensis subsp. wahuensis Kawaihau, Nonou Mts. trail, N exposure; with Acacia koa, Psidium cattleianum, Pandanus tectorius, Gahnia aspera Preserved_Specimen Flynn, T.W. 2780 Kauai BISH 1988-02-26
190 Carex wahuensis subsp. wahuensis Waimea District, Waimea Canyon, north of Kukui Trail relic Acacia-Metrosideros lowland dry to mesic forest dominated by secondary invasive species: primarily Grevillea, Psidium, Lantana, and Hyptis with relictual trees of Myrsine lanaiensis, Dodonaea vicosa, Leptecophylla tameiameiae, Hibiscus waimeae, Diospyros sandwicensis, Pleomele aurea, Sida fallax, Dianella, Gahnia beecheyi Preserved_Specimen terrestrial sedge, relic beneath invasive Lantana camara, occasional Wood, K.R. 15802 Kauai BISH 2014-01-23
191 Carex wahuensis subsp. wahuensis Waimead District. Southeast of Kukui peak. Acacia-Metrosideros lowland dry to mesic forest. With Dodonaea, Hibiscus waimeae, Diospyros, Chrysodracon aurea, Pouteria sandwicensis, Sida fallax, Dianella, Gahnia beecheyi, Leptecophylla, Eragrostis variabilis, Pterdium. Preserved_Specimen Sedge, terrestrial, leaves dark green, not glaucus, in flower, common Wood, K.R. 18358 Kauai BISH 2020-01-21
192 Carex wahuensis subsp. wahuensis Nuuanu Pali Growing sympatrically with Carex wahuensis subsp. herbstii (see Perlman 21101, 21102). Preserved_Specimen Perlman, S. 21103 Oahu BISH 2008-08-06
193 Carex wahuensis subsp. wahuensis Waiahuakua Valley, Hanakoa rocky base of cliff Preserved_Specimen St.John, H. 23142 Kauai BISH 1947-12-29
194 Carex wahuensis subsp. wahuensis West Maui, Lahaina District, Puehuehunui, South side of Kauaula Valley Locally common on ridge in dry shrubland, growing with typical Carex wahuensis, ... Vegetation: Osteomeles anthyllidifolia, Dodonaea viscosa, Bidens menziesii subsp. menziesii, B. micrantha subsp. micrantha, Carex meyenii, Sida fallax, Santalum ellipticum, Chamaesyce celastroides var. amplectens, Wikstroemia oahuensis var. oahuensis, Eragrostis grandis, E. atropioides, Doryopteris decipiens, Pellaea ternifolia, Cocculus orbiculatus. Preserved_Specimen growing with typical C. wahuensis, but differs in the dull and glaucous upper leaf surface (as opposed to deep green and glossy). Oppenheimer, H.L. 20306 Maui BISH 2003-02-17
195 Carex wahuensis subsp. wahuensis Waianae Mts., 1st gulch below Puu Hapapa, 100 yd. above Honouliuli Contour trail wet, shaded; with Cyrtandra, Charpentiera, Urera, ferns Preserved_Specimen Herbst, D.R. 432 Oahu BISH 1967-04-15
196 Carex wahuensis subsp. wahuensis Koolau Divide, opposite Kaluanui ridge, Waimanalo grassy ridge Preserved_Specimen tuft St.John, H. 20182 Oahu BISH 1941-02-16
197 Carex wahuensis subsp. wahuensis Waianae Mts., Puu Hapapa, Honouliuli open ridge Preserved_Specimen St.John, H. 10440 Oahu BISH 1930-03-16
198 Carex wahuensis subsp. wahuensis Waianae Mts., Mt. Kaala, E. Slope, along trail With Acacia koa, Metrosideros Preserved_Specimen Krauss, R.W. 196 Oahu BISH 1948-04-18
199 Carex wahuensis subsp. wahuensis First ridge makai of Mano Divide on the north side of Moanalua Valley. Growing in cracks on an exposed cliff face with Eragrostis, Bidens, Phyllanthus, and Psilotum Preserved_Specimen Herbst, D.R. 1495 Oahu BISH 1970-01-11
200 Carex wahuensis subsp. wahuensis Southwest flank of Mauna Loa, in the ahupuaa of Puu Ohohia Vegetation: Semi closed Metrosideros and Acacia forest, understory Dodonaea, Styphelia, Vaccinium on cinder. Preserved_Specimen Higashino, P.K. 6310 Hawaii BISH 1977-08-05
201 Carex wahuensis subsp. wahuensis Kokee Camp moist woods Preserved_Specimen Degener, O. 8278 Kauai BISH 1926-06-25
202 Carex wahuensis subsp. wahuensis Half way up Kaala from Schofield Fire-Break Trail, Oahu, T.H. Along trail. Preserved_Specimen Degener, O. 17569 Oahu BISH 1936-02-02
203 Carex wahuensis subsp. wahuensis Mauka of Kulani Prison In clearing Preserved_Specimen Degener, O. 21857 Hawaii BISH 1952-02-15
204 Carex wahuensis subsp. wahuensis Kokee, S of main road, N of Waimea Canyon Lookout grove of Pisonia Preserved_Specimen Stone, B.C. 3763 Kauai BISH 1961-08-15
205 Carex wahuensis subsp. wahuensis Koolau Range, Wiliwilinui ridge guava thicket Preserved_Specimen Krauss, R.W. 243 Oahu BISH 1948-06-28
206 Carex wahuensis subsp. wahuensis Waimea, Wainiua trail, Kokee moist woods; with Metrosideros and Osmanthus Preserved_Specimen 2 dm diam. tuft; pergynia green St.John, H. 22880 Kauai BISH 1947-12-21
207 Carex wahuensis subsp. wahuensis W Nonou, Wailua wooded ridge Preserved_Specimen tufts St.John, H. 24925 Kauai BISH 1952-12-31
208 Carex wahuensis subsp. wahuensis Kokee, Awawapuhi trail Acacia koa; steep slope Preserved_Specimen Brenan, J.P.M. 16894 Kauai BISH 1980-03-02
209 Carex wahuensis subsp. wahuensis East Maui, Hana, Auwahi dry forest on aa, rocky Preserved_Specimen Webster, G.L. 13918 Maui BISH 1968-06-25
210 Carex wahuensis subsp. wahuensis Waianae Mts., Honouliuli, E ridge of Puu Kanehoa open woods Preserved_Specimen tufts St.John, H. 14059 Oahu BISH 1934-01-07
211 Carex wahuensis subsp. wahuensis Waianae Mts., trail to Puu Kaua, Dry, N.E. facing slope Preserved_Specimen Krauss, R.W. 126 Oahu BISH 1948-02-29
212 Carex wahuensis subsp. wahuensis Waimea District. Makaha Valley. Diospyros-Metrosideros-Acacia montane mesic forest w/ Diospyros sandwicensis, D. hillebrandii, Alphitonia ponderosa, Pisonia sandwicensis, Dodonaea viscosa, Chrysodracon aurea, Psychotria mariniana, Kadua affinis, Coprosoma waimeae, C. foliosa, Myrsine Preserved_Specimen Sedge, terrestrial, in flower and early fruit, common w/ Carex meyenii. Wood, K.R. 18086 Kauai BISH 2019-02-08
213 Carex wahuensis subsp. wahuensis Koolau Mts., Pupukea, summit trail several Pritchardia stands Preserved_Specimen Stone, B.C. 2789 Oahu BISH 1959-05-04
214 Carex wahuensis subsp. wahuensis Kamoamoa kipuka Closed canopy Diospyros at 8-10m with Erythrina, Santalum, Reynoldsia etc. Rare in kipuka. Preserved_Specimen Stemmermann, R.L. 6925 Hawaii BISH 1985-02-24
215 Carex wahuensis subsp. wahuensis Waimea District. Mohihi Metrosideros polymorpha and Acacia koa montane mesic forest. W/Metrosideros polymorpha, Acacia koa, Dodonaea viscosa, Dianella sandwicensis, Cheirodendron spp., Melicope anisata, Dryopteris wallichiana, Psychotria sp., Wikstroemia oahuensis, Elaeocarpus bifidu Preserved_Specimen Sedge about 3 feet long. Leaves and immature fruit dark green. Mature fruit golden brown Walsh, S. 593 Kauai BISH 2018-06-07
216 Carex wahuensis subsp. wahuensis W Hanakoa Valley dry rocky ridge Preserved_Specimen St.John, H. 23194 Kauai BISH 1947-12-31
217 Carex wahuensis subsp. wahuensis Southwest flank of Mauna Loa, in the ahupuaa of Kapua Vegetation: Open grassy area with Acacia and Metrosideros growing on the perimeter Preserved_Specimen Higashino, P.K. 6117 Hawaii BISH 1977-07-13
218 Carex wahuensis subsp. wahuensis Hill mauka of Olinda on way to Haleakala hill, dry Preserved_Specimen Degener, O. 8254 Maui BISH 1927-06-15
219 Carex wahuensis subsp. wahuensis gulch W of Ualapue wet rocks Preserved_Specimen Degener, O. 8227 Molokai BISH 1928-06-30
220 Carex wahuensis subsp. wahuensis Nuuanu Pali Growing sympatrically with Carex wahuensis subsp. wahuensis (see Perlman 21103). Preserved_Specimen Perlman, S. 21102 Oahu BISH 2008-08-06
221 Carex wahuensis subsp. wahuensis Honokahau Drainage Basin Preserved_Specimen Forbes, C.N. 411 Maui BISH 1917-09-25
222 Carex wahuensis subsp. wahuensis W Maui, Hanaulaiki Preserved_Specimen Broad, blue-green leaves. Growing near # 3734. Hobdy, R.W. 3735 Maui BISH 1995-01-24
223 Carex wahuensis subsp. wahuensis East Maui, NE Haleakala, upper Lauulu trail N of Paliku Cabin with Vaccinium reticulatum, Styphelia, Deschampsia, Holcus, Lythrum Preserved_Specimen Henrickson, J.S. 3574 Maui BISH 1969-06-19
224 Carex wahuensis subsp. wahuensis Bowman trail, ridge W Kalihi Valley wet; with Metrosideros polymorpha, Antidesma, Freycinetia, Scaevola, Hedyotis, Machaerina, Psychotria Preserved_Specimen Perlman, S. 10797 Oahu BISH 1988-02-29
225 Carex wahuensis subsp. wahuensis Kailua, NW of Ulumawao bottom of rocky gully Preserved_Specimen Fosberg, F.R. 13552 Oahu BISH 1937-01-24
226 Carex wahuensis subsp. wahuensis East Maui, Haleakala, 50 m E of Hosmer's Grove rocky ravine, with Metrosideros Preserved_Specimen Higashino, P.K. 619 Maui BISH 1975-06-12
227 Carex wahuensis subsp. wahuensis Kohala Ditch Trail Preserved_Specimen Height: ¼ meters Lee, H.A. 145 Hawaii BISH 1924-10-20
228 Carex wahuensis subsp. wahuensis East Maui, Headwalls of Kipahulu Valley on Mauka ridge. Preserved_Specimen Common name: Oahu sedge Welton, P. 1980 Maui BISH 1997-09-25
229 Carex wahuensis subsp. wahuensis Honopu Beach Preserved_Specimen A sedge Davis, J. Kauai BISH 1984-07-19
230 Carex wahuensis subsp. wahuensis East Maui, Makawao Forest Reserve, Borge Ridge Rd. Preserved_Specimen 3' long stem, perigynium green Hobdy, R.W. 1377 Maui BISH 1982-04-30
231 Carex wahuensis subsp. wahuensis Kanehoa trail Preserved_Specimen Funk, E.J. 93 Oahu BISH 1982-01-15
232 Carex wahuensis subsp. wahuensis W Maui, Hanaulaiki Preserved_Specimen Narrow, bright green leaved plants. Growing near # 3735. Hobdy, R.W. 3734 Maui BISH 1995-01-24
233 Carex wahuensis subsp. wahuensis Makaleha. Ahupuaa of Kamananui. Northern Waianae mountains. Growing on shaded slope with Diospyros sandwicensis, Diospyros hillebrandii, Psidium cattleianum, Schinus terebinthifolius, Aleurites moluccana, Erythrina sandwicensis, Urochloa maxima, Adiantum hispidulum, Salvia coccinea, and others. Preserved_Specimen Thomas, M.K. 1135 Oahu BISH 2025-06-12
234 Carex wahuensis subsp. wahuensis Koolau Range, N-S Halawa ridge Preserved_Specimen Shinohara, B. 210 Oahu BISH 1947-03-02
235 Carex wahuensis subsp. wahuensis Ridge between Niu and Wailupe Preserved_Specimen Forbes, C.N. 2466 Oahu BISH 1917-04-11
236 Carex wahuensis subsp. wahuensis N of Kaala, ridge Preserved_Specimen Degener, O. 17630 Oahu BISH 1937-03-26
237 Carex wahuensis subsp. wahuensis Keawapilau Gulch Preserved_Specimen Welton, P. 637 Oahu BISH 1991-01-20
238 Carex wahuensis subsp. wahuensis Mt. Kaala Preserved_Specimen Bergman, H.F. Oahu BISH 1928-02-11
239 Carex wahuensis subsp. wahuensis Na Pali Trail Preserved_Specimen A Sedge Corn, C. Kauai BISH 1984-07-21
240 Carex wahuensis subsp. wahuensis head of Kuliouou Valley Preserved_Specimen Degener, O. 8272 Oahu BISH 1929-12-11
241 Carex wahuensis subsp. wahuensis Keopukai, Maui Preserved_Specimen Rare to occasional. Scattered clumps. Starr, F. 5040530 Maui BISH 2005-04-05
242 Carex wahuensis subsp. wahuensis Kalaupapa; Papapaiki, outside fence Preserved_Specimen rhizomatous perennial Wysong, M.L. 574 Molokai BISH 2005-01-11
243 Carex wahuensis subsp. wahuensis S of Ekahanui Preserved_Specimen varigated Degener, O. 24349 Oahu BISH 1936-04-22
244 Carex wahuensis subsp. wahuensis East Maui, N Haleakala, 1.75 mi N of Paliku Cabin, below Puu Alaea with Metrosideros, Vaccinium, Styphelia, Cheirodendron Preserved_Specimen Henrickson, J.S. 3698 Maui BISH 1969-06-22
245 Carex wahuensis subsp. wahuensis Palikea Gulch, central ridge steep slopes, open Preserved_Specimen Takeuchi, W.N. 72 Oahu BISH 1984-02-20
246 Carex wahuensis subsp. wahuensis Olokele ditch trail, Makaweli shaded rocky hillside Preserved_Specimen St.John, H. 23078 Kauai BISH 1947-12-26
247 Carex wahuensis subsp. wahuensis Haleakala, E of rest house scrub Preserved_Specimen Degener, O. 12552 Maui BISH 1939-07-26
248 Carex wahuensis subsp. wahuensis N Haupu base of cliff Preserved_Specimen MacDaniels, L.H. 723 Kauai BISH 1927-02-16
249 Carex wahuensis subsp. wahuensis Kaumokunui Gulch, Mokuleia Preserved_Specimen Degener, O. 17611 Oahu BISH 1936-04-13
250 Carex wahuensis subsp. wahuensis Preserved_Specimen Degener, O. 5263 BISH 1927-02-06
251 Carex wahuensis subsp. wahuensis East Maui, Makaiwa gulch Preserved_Specimen Rock, J.F.C. 8747 Maui BISH 1911-04-01
252 Carex wahuensis subsp. wahuensis N of Kaala Preserved_Specimen Degener, O. 17610 Oahu BISH 1937-04-26
253 Carex wahuensis subsp. wahuensis Koolau Mts., Pia Valley, ~2 mi into gulch Preserved_Specimen Sohmer, S.H. 6473 Oahu BISH 1970-06-10
254 Carex wahuensis subsp. wahuensis Between Palehua and Palikea Preserved_Specimen Degener, O. 17612 Oahu BISH 1935-12-16
255 Carex wahuensis subsp. wahuensis Waianae Mts., central Lualualei, below Kanehoa, between valleys 1 and 2 Preserved_Specimen Christophersen, E. 3695 Oahu BISH 1932-06-01
256 Carex wahuensis subsp. wahuensis East Maui, Lower Shelf of Kipahulu Valley on streambank near Delta Camp. Preserved_Specimen Welton, P. 1631 Maui BISH 1992-06-04
257 Carex wahuensis subsp. wahuensis East Maui, Makawao Preserved_Specimen Rock, J.F.C. 8747 Maui BISH 1911-04-01
258 Carex wahuensis subsp. wahuensis East Maui, Hana District, Auwahi Preserved_Specimen Hobdy, R.W. 1731 Maui BISH 1983-01-21
259 Carex wahuensis subsp. wahuensis Kaiholena Preserved_Specimen Munro, G.C. 497 Lanai BISH 1914-02-17
260 Carex wahuensis subsp. wahuensis E Palikea, trail to Pohakea Pass Preserved_Specimen Degener, O. 17437 Oahu BISH 1936-03-01
261 Carex wahuensis subsp. wahuensis Puu Kaua trail Preserved_Specimen Woolford, H.A. 232 Oahu BISH 1955-12-11
262 Carex wahuensis subsp. wahuensis Kaiholena Preserved_Specimen Munro, G.C. 297 Lanai BISH 1914-02-17
263 Carex wahuensis subsp. wahuensis Waianae Mts., Dupont Trail, opposite Kaupakuhale Preserved_Specimen Krajina, V.J. 620225313 Oahu BISH 1962-02-25
264 Carex wahuensis subsp. wahuensis Pig God trail, Punaluu, near top Preserved_Specimen Degener, O. 17629 Oahu BISH 1937-03-29
265 Carex wahuensis subsp. wahuensis Haleakala Preserved_Specimen Degener, O. 8262 Maui BISH 1927-08-14
266 Carex wahuensis subsp. wahuensis Haleakala, Puu Pani Preserved_Specimen Forbes, C.N. 1887 Maui BISH 1920-03-08
267 Carex wahuensis subsp. wahuensis West Maui, Wailuku, Iao Valley, above Stream Preserved_Specimen Wagner, W.L. 5831 Maui BISH 1988-03-06
268 Carex wahuensis subsp. wahuensis Napali coast, Hanakapiai, Maunapuluo Preserved_Specimen St.John, H. 10893 Kauai BISH 1930-12-30
269 Carex wahuensis subsp. wahuensis Waianae Mts., Mauna Kapu Preserved_Specimen Webster, G.L. 1278 Oahu BISH 1948-02-04
270 Carex wahuensis subsp. wahuensis Waianae Mts., 1/4 mi. N of Puu Kaua, lateral E ridge Preserved_Specimen Cowan, R.S. 851 Oahu BISH 1948-02-29
271 Carex wahuensis subsp. wahuensis Kalualea, Koloa Preserved_Specimen Degener, O. 17589 Kauai BISH 1939-12-31
272 Carex wahuensis subsp. wahuensis Nuu, Waiopaa ranch Preserved_Specimen Forbes, C.N. 1862 Maui BISH 1920-03-06
273 Carex wahuensis subsp. wahuensis Waialae Iki Preserved_Specimen Forbes, C.N. 2425 Oahu BISH 1917-02-28
274 Carex wahuensis subsp. wahuensis W Waimea drainage basin Preserved_Specimen Forbes, C.N. 1096 Kauai BISH 1917-07-03
275 Carex wahuensis subsp. wahuensis Waialae Valley Preserved_Specimen Forbes, C.N. 2518 Oahu BISH 1919-01-08
276 Carex wahuensis subsp. wahuensis Mahanaloa Valley, N facing slope trail Preserved_Specimen Flynn, T.W. 665 Kauai BISH 1983-11-29
277 Carex wahuensis subsp. wahuensis E of Kokee, forest road Preserved_Specimen Alexander, A.M. 5210 Kauai BISH 1947-03-08
278 Carex wahuensis subsp. wahuensis East Maui, Nuu, Waiopaa Preserved_Specimen Forbes, C.N. 1862 Maui BISH 1920-03-06
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